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Gabor Medal20epigenetics30Buchanan Medal37Shaw Prize in Life Science and Medicine41Portland Press Excellence in Science Award42The Brain Prize43Grand Prix Charles-Leopold Mayer50Fellow of the AACR Academy73Louis-Jeantet Prize for Medicine96BBVA Foundation Frontiers of Knowledge Award129Canada Gairdner International Award353Clarivate Citation Laureates400genetics416Fellow of the Academy of Medical Sciences887Fellow of the Royal Society of Edinburgh2,286Fellow of the Royal Society5,768Knight Bachelor6,416Commander of the Order of the British Empire6,422194716,350United Kingdom142,5281940143,130English435,916201,347,152male1,587,100