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Vanderbilt Prize in Biomedical Science15Gruber Prize in Genetics27FASEB Excellence in Science Award30William C. Rose Award39Wolf Prize in Medicine66Warren Alpert Foundation Prize77Canada Gairdner International Award353biochemistry751Fellow of the American Association for the Advancement of Science4,535Doctor of Philosophy6,244195215,8621950161,666female412,920English435,916United States442,023201,347,152